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in reply to Re^2: Extracting values from nested hashrefs
in thread Extracting values from nested hashrefs

You can use
map $_->{humps}, values %$animals;
If your structure becomes even more complicated (e.g. different level of nesting), you should use recursion (or CPAN).
لսႽ† ᥲᥒ⚪⟊Ⴙᘓᖇ Ꮅᘓᖇ⎱ Ⴙᥲ𝇋ƙᘓᖇ