use Regexp::Common; my $balanced = qr/[^()]+|$RE{balanced}{-parens=>'()'}/; sub extract { my ($want, $from) = @_; my $nested = qr/$balanced* \Q$want\E $balanced*/x; $from =~ m/( \( $nested \) | \Q$want\E )/x; return $1; } my $expr = '(((B,G),A,(A,B)),C,(D,(E))),F'; for ('A'..'H') { print "$_: ", extract($_,$expr),"\n"; }