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use strict;
use warnings;
...
        return '*';
    }
}

>M01096:4:000000000-A23M1:1:1101:15974:1529
VTECGWQTAGCRMY
...
LSHCDVKDWMCWLL
>M01096:4:000000000-A23M1:1:1101:13838:1554
AWTCVEIDGHFSMN